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Biography
other usesTaxobox| fossil_range = fossil range|Valanginian|Recent| image = Lobster NSRW.jpg| image_caption = American lobster , Homarus americanus | regnum = Animal ia| phylum = Arthropod a| subphylum = Crustacean|Crustacea | classis = Malacostraca | ordo = Decapoda | subordo = Pleocyemata | infraordo = Astacidea | familia = Nephropidae | familia_authority = James Dwight Dana|Dana , 1852| subdivision_ranks = Genus|Genera & nbsp;cite journal |journal = Raffles Bulletin of Zoology |year=2009 |volume = Suppl. 21 |pages=1–109 |title = A classification of living and fossil genera of decapod crustaceans |author=Sammy De Grave, N. Dean Pentcheff, Shane T. Ahyong et al. |url= http://rmbr.nus.edu.sg/rbz/biblio/s21/s21rbz1-109.pdf| subdivision =
Acanthacaris Bate, 1888
Eunephrops Smith, 1885
Cape lobster|Homarinus Kornfield, Williams & Steneck, 1995
Homarus Weber, 1795
Hoploparia † M’Coy, 1849
Jagtia † Tshudy & Sorhannus, 2000
Metanephrops Jenkins, 1972
Nephropides Manning, 1969
Nephrops Leach, 1814
Nephropsis Wood-Mason, 1873
Oncopareia † Bosquet, 1854
Palaeonephrops † Mertin, 1941
Paraclythia † Fritsch & Kafka, 1887
Pseudohomarus † van Hoepen, 1962
Thaumastocheles Wood-Mason, 1874
Thaumastochelopsis Bruce, 1988
Thymopides Burukovsky & Averin, 1977
Thymops Holthuis, 1974
Thymopsis nilenta|Thymopsis Holthuis, 1974
Clawed lobsters comprise a family (biology)|family ( Nephropidae , sometimes also Homaridae ) of large marine crustacean s. Highly prized as seafood , lobsters are economically important, and are often one of the most profitable commodities in coastal areas they populate.cite web |url= http://www.geog.mcgill.ca/climatechange/ReportsMap/lobsterRpt.pdf |title= Homarus americanus , American lobster |date=27 June 2007 |publisher= McGill University
Though several groups of crustaceans are known as lobsters, the clawed lobsters are most often associated with the name. Clawed lobsters are not closely related to spiny lobster s or slipper lobster s, which have no claws ( Chela (organ)|chelae ), or squat lobster s. The closest relatives of clawed lobsters are the reef lobster s and the three families of freshwater crayfish .
Evolution
The fossil record of clawed lobsters extends back at least to the Valanginian|Valanginian Age of the Cretaceous .cite journal |journal= Journal of Paleontology |year=2005 |volume=79 |issue=5 |pages=961–968 |doi=10.1666/0022-3360(2005)0790961:HAANSO2.0.CO;2 |title= Hoploparia albertaensis , a new species of clawed lobster (Nephropidae) from the Late Coniacean, shallow-marine Bad Heart Formation of northwestern Alberta, Canada |author=Dale Tshudy, W. Steven Donaldson, Christopher Collom, Rodney M. Feldmann & Carrie E. Schweitzer
Description
Lobsters are invertebrate s with a hard protective exoskeleton . Like most arthropod s, lobsters must ecdysis|molt in order to grow, which leaves them vulnerable. During the molting process, several species change color. Lobsters have 10 walking legs; the front three pairs bear claws, the first of which are larger than the others.cite book |editor=Mark W. Denny & Steven Dean Gaines |year=2007 |title=Encyclopedia of tidepools and rocky shores |publisher= University of California Press |isbn=978-0-520-25118-2 |author=Carlos Robles |chapter=Lobsters |pages=333–335 |url= http://books.google.co.uk/books? id=uufQnE7MzMkC& pg=PA333 Although, like most other arthropods, lobsters are largely bilateral symmetry|bilaterally symmetrical , they often possess unequal, specialized claws, like the king crab .
Lobster anatomy includes the cephalothorax which fuses the head and the thorax , both of which are covered by a chitin ous carapace , and the abdomen. The lobster's head bears Antenna (biology)|antennae , antennules, Mandible (arthropod)|mandibles , the first and second Maxilla (arthropod)|maxillae , and the first, second, and third Decapod anatomy#Thorax / Pereon|maxillipeds . Because lobsters live in a murky environment at the bottom of the ocean, they mostly use their antennae as sensors. The lobster eye has a reflective structure above a convex retina. In contrast, most complex eyes use refractive ray concentrators (lenses) and a concave retina.cite journal |author=M. F. Land |title=Superposition images are formed by reflection in the eyes of some oceanic decapod Crustacea |journal= Nature (journal)|Nature |year=1976 |volume=263 |pages=764–765 |doi=10.1038/263764a0 |pmid=995187 |issue=5580 The abdomen includes decapod anatomy|swimmerets and its tail is composed of Decapod anatomy#Abdomen / Pleon|uropods and the telson .
Lobsters, like snails and spiders, have blue blood due to the presence of haemocyanin which contains copper cite web |url= http://resources.schoolscience.co.uk/cda/11-14/biology/copch31pg1.html |title=Copper for life – Vital copper |publisher= Association for Science Education (in contrast, vertebrates and many other animals have red blood from iron -rich haemoglobin ). Lobsters possess a green hepatopancreas , called the tomalley by chefs, which functions as the animal's liver and pancreas .cite journal |url= http://www.rsc.org/delivery/_ArticleLinking/DisplayArticleForFree.cfm? doi=b314101b& JournalCode=JA |doi=10.1039/b314101b |title=Arsenic speciation in marine certified reference materials |year=2004 |author=Shona Mcsheehy & Zoltán Mester |journal= Journal of Analytical Atomic Spectrometry |volume=19 |pages=373–380 |issue=3
In general, lobsters are convert|25|-|50|cm|0|in long, and move by slowly walking on the sea floor. However, when they flee, they swim backwards quickly by curling and uncurling their abdomen . A speed of convert|5|m/s|mph has been recorded.cite web |url= http://www.osl.gc.ca/homard/en/faq.html |title=The American lobster – frequently asked questions |publisher=St. Lawrence Observatory, Fisheries and Oceans Canada |date=October 19, 2005 This is known as the caridoid escape reaction .
Longevity
Recent research suggests that lobsters may not slow down, weaken, or lose fertility with age. In fact, older lobsters are more fertile than younger lobsters. This longevity may be due to telomerase , an enzyme that repairs DNA sequence s of the form "TTAGGG."cite web |url= http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/T/Telomeres.html |author=John W. Kimball |date=November 25, 2008 |title=Telomeres This sequence, repeated hundreds of times, occurs at the ends of chromosomes, which are referred to as the telomeres .cite web |url= http://animals.howstuffworks.com/marine-life/400-pound-lobster.htm/printable |title=Is there a 400 pound lobster out there? |author=Jacob Silverman |publisher= howstuffworks cite book |title= Consider the Lobster|Consider the Lobster and Other Essays |author=David Foster Wallace |publisher= Little, Brown & Company |year=2005 |isbn=0-316-15611-6 |chapter=Consider the Lobster |chapterurl= http://www.gourmet.com/magazine/2000s/2004/08/consider_the_lobster? currentPage=1 |authorlink=David Foster Wallace It has been argued that lobsters may exhibit negligible senescence and some scientists have claimed that they could effectively live indefinitely, barring injury, disease, capture, etc.cite journal |author=John C. Guerin |journal= Annals of the New York Academy of Sciences |volume=1019 |issue=1 |pages=518–520 |url= http://www.blackwell-synergy.com/doi/abs/10.1196/annals.1297.096 |doi=10.1196/annals.1297.096 |pmid=15247078 |year=2006 |title=Emerging area of aging research: long-lived animals with "negligible senescence" Their longevity allows them to reach impressive sizes. According to the Guinness World Records , the largest lobster was caught in Nova Scotia , Canada , and weighed convert|20.15|kg|lb.cite web|url= http://www.guinnessworldrecords.com/content_pages/record.asp? recordid=51451 |publisher= Guinness World Records |title=Heaviest marine crustacean |accessdate=August 3, 2006 |archiveurl= http://web.archive.org/web/20060528192250/ http://www.guinnessworldrecords.com/content_pages/record.asp? recordid=51451 |archivedate=May 28, 2006cite news |url= http://news.bbc.co.uk/1/hi/england/kent/5118370.stm |title=Giant lobster landed by boy, 16 |date=June 26, 2006 |publisher= BBC News
Symbion
Main|SymbionAnimals of the genus Symbion , the only member of the animal phylum Cycliophora, live exclusively on lobster gill s and mouthparts.cite journal |author=M. Obst, P. Funch & G. Giribet |year=2005 |title= Hidden diversity and host specificity in cycliophorans: a phylogeographic analysis along the North Atlantic and Mediterranean Sea |journal= Molecular Ecology (journal)|Molecular Ecology |pmid=16313603 |volume=14 |issue=14 |pages=4427–4440 |url= http://www.blackwell-synergy.com/doi/abs/10.1111/j.1365-294X.2005.02752.x |doi=10.1111/j.1365-294X.2005.02752.x
Ecology
Lobsters are found in all oceans. They live on rocky, sandy, or muddy bottoms from the shoreline to beyond the edge of the continental shelf . They generally live singly in crevices or in burrows under rocks.
Lobsters are omnivores and typically eat live prey such as fish, mollusks, other crustaceans, worms, and some plant life. They scavenge if necessary, and may resort to Cannibalism (zoology)|cannibalism in captivity; however, this has not been observed in the wild. Although lobster skin has been found in lobster stomachs, this is because lobsters eat their shed skin after ecdysis|molting .cite web |url= http://marinebio.org/species.asp? id=533 |title= Homarus americanus , Atlantic lobster |accessdate=December 27, 2006 |publisher=MarineBio.org
Gastronomy
redirect|Lobster claw|the species of flowering plants|Lobster-clawnutritionalvalue | name=Lobster | kJ=410| protein=20.5 g | fat=0.59 g | satfat=0.107 g | monofat=0.091 g | polyfat=0.16 g | carbs=0 g | fibre=0 g | | sugars=0 g | iron_mg=2 | calcium_mg=6 | magnesium_mg=8 | phosphorus_mg=15 | potassium_mg=0 | zinc_mg=15 | vitC_mg=0 | pantothenic_mg=2 | vitB6_mg=4 | folate_ug=2 | thiamin_mg=0 | riboflavin_mg=4 | niacin_mg=4 | right=1 | source_usda=1 stack||float=left Lobster recipes include Lobster Newberg and Lobster Thermidor . Lobster is used in soup, bisque (food)|bisque , lobster roll s, and cappon magro . Lobster meat may be dipped in clarified butter , resulting in a sweetened flavour.
Cooks boil or steam live lobsters. The lobster cooks for seven minutes for the first pound and three minutes for each additional pound.cite web |url= http://www.atwoodlobster.com/site/cookinglobster.asp |title=Cooking lobsters |publisher=Atwood Lobster Company |accessdate=June 30, 2007 |archiveurl = http://web.archive.org/web/20070607114042/ http://www.atwoodlobster.com/site/cookinglobster.asp |archivedate = June 7, 2007
According to the United States Food and Drug Administration (FDA), the mean level of mercury (element)|mercury in American lobster is 0.31& nbsp; parts per million|ppm .cite web |url= http://www.fda.gov/Food/FoodSafety/Product-SpecificInformation/Seafood/FoodbornePathogensContaminants/Methylmercury/ucm115644.htm |title=Mercury Levels in Commercial Fish and Shellfish |publisher= Food and Drug Administration |accessdate=December 25, 2009
History
In North America, the American lobster did not achieve popularity until the mid-19th century, when New Yorkers and Bostonians developed a taste for it, and commercial lobster fisheries only flourished after the development of the lobster smack .cite book |author=Colin Woodard |url= http://www.colinwoodard.com/lobstercoast |title=The Lobster Coast |publisher=New York: Viking/Penguin |isbn=0-670-03324-3 |year=2004 |pages=170–180 Prior to this time, lobster was considered a mark of poverty or as a food for indentured servants or lower members of society in Maine , Massachusetts and the Canadian Maritimes , and servants specified in employment agreements that they would not eat lobster more than twice per week.cite news |url= http://www.timesonline.co.uk/tol/news/world/us_and_americas/article581926.ece |title=How lobster went up in the world |publisher= The Times |date=October 24, 2005 | location=London | first=Mark | last=Henderson | accessdate=May 11, 2010 American lobster was initially deemed worthy only of being used as fertilizer or fish bait, and it was not until well into the twentieth century that it was viewed as more than a low-priced canned staple food.cite book|first=Paul |last=Johnson |title=Fish Forever: The Definitive Guide to Understanding, Selecting, and Preparing Healthy, Delicious, and Environmentally Sustainable Seafood | chapter=Lobster | publisher= John Wiley & Sons |year=2007 |isbn=978-0-7645-8779-5|pages=163–175
Caught lobsters are graded as new-shell, hard-shell and old-shell and, because lobsters that have recently shed their shells are the most delicate, there is an inverse relationship between the price of American lobster and its flavor. New-shell lobsters have paper-thin shells and a worse meat-to-shell ratio, but what meat exists is very sweet. However, the lobsters are so delicate that even transport to Boston almost kills them, making the market for new-shell lobsters strictly local to the fishing towns where they are offloaded. Hard-shell lobsters with firm shells but with less sweet meat can survive shipping to Boston, New York and even Los Angeles so they command a higher price than new-shell lobsters. Meanwhile, old-shell lobsters, which have not shed since the previous season and have a coarser flavor, can be air-shipped anywhere in the world and arrive alive, making them the most expensive. One seafood guide notes that an eight dollar lobster dinner at a restaurant overlooking fishing piers in Maine is consistently delicious, while "the eighty-dollar lobster in a three-star Paris restaurant is apt to be as much about presentation as flavor".
Animal welfare issues
See|Pain in crustaceansThe most common way of killing a lobster is by placing it live in boiling water (with or without spending a period of time in a freezer) or by splitting it by severing the body in half lengthwise. Lobsters may also be killed or rendered insensate immediately before boiling through a stab into the brain, in the belief that this will stop suffering. However, a lobster's brain operates from not one but several Ganglion|ganglia and disabling only the frontal ganglion does not usually result in death or unconsciousness. The boiling method is illegal in some places, such as in Reggio Emilia , Italy, where offenders face fines of up to Euro|€ 495.cite news |url= http://www.telegraph.co.uk/news/worldnews/europe/italy/1456270/Italian-animal-rights-law-puts-lobster-off-the-menu.html |title=Italian animal rights law puts lobster off the menu |author=Bruce Johnston |date=March 6, 2004 |publisher= The Daily Telegraph|Daily Telegraph | location=London
Fishery and aquaculture
main|lobster fishingLobsters are caught using lobster trap|baited, one-way traps with a color-coded marker buoy to mark cages. Lobster is fished in water between convert|1|and|500|fathom|m|sigfig=1, although some lobsters live at convert|2000|fathom|m. Cages are of plastic-coated galvanized steel or wood. A lobster fisher may tend as many as 2,000 traps. Around the year 2000, due to overfishing and high demand, lobster aquaculture expanded.cite web |url= http://articles.uwphoto.no/articles_folder/lobster_farming_in_Norway.htm |author=Asbjørn Drengstig, Tormod Drengstig & Tore S. Kristiansen |publisher=UWPhoto ANS |title=Recent development on lobster farming in Norway – prospects and possibilities As of 2008, no lobster aquaculture operation had achieved commercial success, due mainly to the fact that lobsters eat each other (cannibalism) and the slow growth of the species; these two problems make it difficult to make lobster aquaculture profitable. http://www.gma.org/lobsters/trivia.html Problems surrounding lobster aquaculture The New England Aquarium nonetheless maintains a year-round production facility of American lobster, and further research is being conducted. http://www.neaq.org/conservation_and_research/projects/fisheries_bycatch_aquaculture/aquaculture_research/american_lobster_aquaculture_research/background/neaq_lobster_lab.php Lobster lab
Species
Multiple image| width = 220 | header = Examples of Nephropidae | direction = vertical | align = right | image2 = Metanephrops japonicus edit.jpg | caption2 = Metanephrops japonicus | image3 = Nephropsis rosea.jpg | caption3 = Nephropsis rosea | image1 = Acanthocaris tenuimana.jpg | caption1 = Acanthacaris tenuimana This list contains all extant species in the family Nephropidae:cite journal |author=Tin-Yam Chan |year=2010 |chapter=Annotated checklist of the world's marine lobsters (Crustacea: Decapoda: Astacidea, Glypheidea, Achelata, Polychelida) |editor=Martyn E. Y. Low and S. H. Tan |title=Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) and marine lobsters of the world |journal= Zootaxa |volume=Suppl. 23 |pages=153–181 |url= http://rmbr.nus.edu.sg/rbz/biblio/s23/s23rbz153-181.pdf |format= Portable Document Format|PDF div col|colwidth=30em
Cite book |year=1991 |url= http://nlbif.eti.uva.nl/bis/lobsters.php? menuentry=inleiding |title=Marine Lobsters of the World |author=Lipke Holthuis |publisher = Food and Agriculture Organization |authorlink= Lipke Holthuis
http://www.lobsterscience.ca/ Atlantic Veterinary College Lobster Science Centre