Racoon

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Biography

The raccoon , Procyon lotor (sometimes spelled racoon ), also known as the common raccoon ,Zeveloff, p. 42 North American raccoon ,Zeveloff, p. 1 northern raccoon and colloquially as coon ,Zeveloff, p. 2 is a medium-sized mammal native to North America. It is the largest of the procyonidae|procyonid family, having a body length of and a body weight of . The raccoon is usually nocturnality|nocturnal and is omnivore|omnivorous , with a diet consisting of about 40% invertebrate s, 33% plant foods, and 27% vertebrate s. It has a grayish coat, of which almost 90% is dense underfur , which insulates against cold weather. Two of its most distinctive features are its extremely dexterous front paw s and its Domino mask|facial mask , which are themes in the Native American mythology|mythology of several Native American tribes . Raccoons are noted for their intelligence, with studies showing that they are able to remember the solution to tasks up to three years later.

The original habitat s of the raccoon are deciduous and Temperate broadleaf and mixed forests|mixed forests of North America, but due to their adaptability they have extended their range to mountainous areas, Salt marsh|coastal marshes , and urban area s, where many homeowners consider them to be pests. As a result of escapes and deliberate introduced species|introductions in the mid-20th century, raccoons are now also distributed across the European mainland, the Caucasus region and Japan.

Though previously thought to be solitary, there is now evidence that raccoons engage in gender-specific social behavior . Related females often share a common area, while unrelated males live together in groups of up to four animals to maintain their positions against foreign males during the mating season, and other potential invaders. Home range sizes vary anywhere from 3 hectares for females in cities to 50& nbsp;km² for males in prairie s (7 acres to 20& nbsp;sq& nbsp;mi). After a gestation|gestation period of about 65 days, two to five young, known as "kits", are born in spring. The kits are subsequently raised by their mother until dispersion in late fall. Although captive raccoons have been known to live over 20 years, their average life expectancy in the wild is only 1.8 to 3.1& nbsp;years. In many areas hunting and vehicular injury are the two most common causes of death.

Etymology

The word "raccoon" was adopted into English from the native Powhatan language|Powhatan term, as used in the Virginia Colony . It was recorded on Captain John Smith 's list of Powhatan words as aroughcun , and on that of William Strachey as arathkone . It has also been identified as a Proto-Algonquian root *ahrah-koon-em , meaning " one who rubs, scrubs and scratches with its hands".Holmgren, p. 23; Zeveloff, p. 2

Similarly, Spanish colonization of the Americas|Spanish colonists adopted the Spanish word from the Nahuatl mapachitli of the Aztec s, meaning " one who takes everything in its hands".Holmgren, p. 52 In many languages, the raccoon is named for its characteristic dousing behavior in conjunction with that language's term for bear , for example in German, in Italian, in Hungarian language|Hungarian and araiguma () in Japanese. In French and Portuguese (in Portugal), the washing behavior is combined with these languages' term for rat , yielding, respectively, and ratão-lavadeiro .

The colloquialism|colloquial abbreviation coon is used in words like coonskin for fur clothing and in phrases like old coon as a self-designation of animal trapping|trappers .Holmgren, pp. 75–76; Zeveloff, p. 2 However, the clipped form is also in use as an Ethnic slur#C|ethnic slur . The raccoon's scientific name, Procyon lotor , is New Latin|neo-Latin , meaning "before-dog washer", with lotor Latin for "washer" and Procyon Latinized Greek language|Greek from p??-, "before" and ????, "dog".

Taxonomy

In the first decades after its discovery by the members of the expedition of Christopher Columbus – the first person to leave a written record about the species – taxonomy|taxonomists thought the raccoon was related to many different species, including Canidae|dogs , Felidae|cats , badger s and particularly bear s.Holmgren, pp. 47–67 Carl Linnaeus , the father of modern taxonomy, placed the raccoon in the genus Ursus (genus)|Ursus , first as Ursus cauda elongata ("long-tailed bear") in the second edition of his Systema Naturae (1740), then as Ursus Lotor ("washer bear") in the tenth edition (1758–59).Holmgren, pp. 64–67; Zeveloff, pp. 4–6 In 1780, Gottlieb Conrad Christian Storr placed the raccoon in its own genus Procyon (genus)|Procyon , which can be translated as either "before the dog" or "doglike".Holmgren, pp. 68–69; Zeveloff, p. 6 It is also possible that Storr had its nocturnality|nocturnal lifestyle in mind and chose the star Procyon as eponym for the species.Hohmann, p. 44; Holmgren, p. 68

Evolution

Based on fossil evidence from France and Germany, the first known members of the family Procyonidae lived in Europe in the late Oligocene about 25 million years ago.Zeveloff, p. 19 Similar tooth and skull structures suggest procyonids and Mustelidae|weasels share a common ancestor, but molecular analysis indicates a closer relationship between raccoons and bears.Zeveloff, pp. 16–18, 26 After the then-existing species crossed the Bering Strait at least six million years later, the center of its distribution was probably in Central America.Zeveloff, pp. 20, 23 Coati s ( Nasua and Nasuella ) and raccoons ( Procyon ) have been considered to possibly share common descent from a species in the genus Paranasua present between 5.2 and 6.0& nbsp;million years ago.Zeveloff, p. 24 This assumption, based on morphological comparisons, conflicts with a 2006 genetic analysis which indicates raccoons are more closely related to Bassariscus|ringtails . Unlike other procyonids, such as the crab-eating raccoon ( Procyon cancrivorus ), the ancestors of the common raccoon left tropics|tropical and subtropics|subtropical areas and migrated farther north about 4& nbsp;million years ago, in a migration that has been confirmed by the discovery in the Great Plains of fossils dating back to the middle of the Pliocene .Hohmann, p. 46; Zeveloff, p. 24

Subspecies

Four subspecies of raccoon Endemism|endemic to small Central American and Caribbean islands were often regarded as distinct species after their discovery. These are the Bahaman raccoon and Guadeloupe raccoon , which are very similar to each other; the Tres Marias raccoon , which is larger than average and has an angular skull; and the extinction|extinct Barbados raccoon . Studies of their morphological and genetic traits in 1999, 2003 and 2005 led all these island raccoon s to be listed as subspecies of the common raccoon in the third edition of Mammal Species of the World (2005).Zeveloff, pp. 42–46 A fifth island raccoon population, the Cozumel raccoon , which weighs only and has notably small teeth, is still regarded as a separate species.

The four smallest raccoon subspecies, with an average weight of , are found along the southern coast of Florida and on the adjacent islands; an example is the Ten Thousand Island raccoon ( Procyon lotor marinus ).Zeveloff, pp. 59, 82–83 Most of the other 15 subspecies differ only slightly from each other in coat color, size and other physical characteristics.MacClintock, p. 9; Zeveloff, pp. 79–89 The two most widespread subspecies are the eastern raccoon ( Procyon lotor lotor ) and the upper Mississippi Valley raccoon ( Procyon lotor hirtus ). Both share a comparatively dark coat with long hairs, but the upper Mississippi Valley raccoon is larger than the eastern raccoon. The eastern raccoon occurs in all US states and Canadian provinces to the north of South Carolina and Tennessee . The adjacent range of the upper Mississippi Valley raccoon covers all US states and Canadian provinces to the north of Louisiana , Texas and New Mexico .Zeveloff, pp. 79–81, 84

Description

Physical characteristics

Head to hindquarters, raccoons measure between , not including the bushy tail which can measure between , but is usually not much longer than .Hohmann, p. 77; Lagoni-Hansen, p. 15; Zeveloff, p. 58 The shoulder height is between .Lagoni-Hansen, p. 16 The skull of the adult male measures 94.3–125.8& nbsp;mm long and 60.2–89.1& nbsp;mm wide, while that of the female measures 89.4–115.9& nbsp;mm long and 58.3–81.2& nbsp;mm wide. The body weight of an adult raccoon varies considerably with habitat ; it can range from , but is usually between . The smallest specimens are found in Southern Florida, while those near the northern limits of the raccoon's range tend to be the largest (see Bergmann's rule ).Zeveloff, pp. 58–59 Males are usually 15 to 20% heavier than females.Lagoni-Hansen, p. 18 At the beginning of winter, a raccoon can weigh twice as much as in spring because of fat storage.Hohmann, p.47–48; MacClintock, p. 44; Zeveloff, p. 108 It is one of the most variably sized of all mammals. The largest recorded wild raccoon weighed and measured in total length, by far the largest size recorded for a procyonid.MacClintock, p. 8; Zeveloff, p. 59Boitani, Luigi, ''Simon & Schuster's Guide to Mammals . Simon & Schuster/Touchstone Books (1984), ISBN 978-0-671-42805-1

The most characteristic physical feature of the raccoon is the area of black fur around the eyes, which contrasts sharply with the surrounding white face coloring. This is reminiscent of a " Domino mask|bandit's mask " and has thus enhanced the animal's reputation for mischief.Bartussek, p.6; Zeveloff, p. 61 The slightly rounded ears are also bordered by white fur. Raccoons are assumed to recognize the facial expression and posture of other members of their species more quickly because of the conspicuous facial coloration and the alternating light and dark rings on the tail. The rings resemble those of a ringtail lemur .Hohmann, pp. 65–66 The dark mask may also reduce glare (vision)|glare and thus enhance night vision .MacClintock, pp. 5–6; Zeveloff, p. 63 On other parts of the body, the long and stiff guard hair s, which shed moisture, are usually colored in shades of gray and, to a lesser extent, brown.Zeveloff, p. 60 Raccoons with a very dark coat are more common in the German population because individuals with such coloring were among those initially released to the wild. The dense underfur , which accounts for almost 90% of the coat, insulates against cold weather and is composed of long hairs.

The raccoon, whose method of Terrestrial locomotion|locomotion is usually considered to be plantigrade , can stand on its hind legs to examine objects with its front paws.Hohmann, p. 57; Zeveloff, p. 71–72 As raccoons have short legs compared to their compact torso, they are usually not able either to run quickly or jump great distances.Hohmann, p. 93; Zeveloff, p. 72 Their top speed over short distances is .MacClintock, p. 28 Raccoons can swim with an average speed of about and can stay in the water for several hours.MacClintock, p. 33; Zeveloff, p. 72 For climbing down a tree headfirst—an unusual ability for a mammal of its size—a raccoon rotates its hind feet so they are pointing backwards.MacClintock, p. 30; Zeveloff, p. 72 Raccoons have a dual cooling system to Thermoregulation|regulate their temperature ; that is, they are able to both sweat and pant for heat dissipation.MacClintock, p. 29; Zeveloff, p. 73

Raccoon skulls have a short and wide facial region and a voluminous braincase . The Facial skeleton|facial length of the skull is less than the Neurocranium|cranial , and their nasal bone s are short and quite broad. The auditory bulla e are inflated in form, and the sagittal crest is weakly developed. The dentition — 40 teeth with the dental formula: — is adapted to their omnivore|omnivorous diet: the carnassial s are not as sharp and pointed as those of a full-time carnivore , but the molar (tooth)|molars are not as wide as those of a herbivory|herbivore .Zeveloff, p. 64 The baculum|penis bone of males is about long and strongly bent at the front end and is often used by biologists to classify reproductive status of specimens.Hohmann, p. 27; MacClintock, p. 84 Seven of the thirteen identified vocal calls are used in Animal communication|communication between the mother and her kits, one of these being the birdlike twittering of newborns.Hohmann, p. 66; MacClintock, p. 92; Zeveloff, p. 73

Senses

The most important sense for the raccoon is its Somatosensory system|sense of touch .Bartussek, p. 13; Hohmann, p. 55; Zeveloff, p. 70 The "hyper sensitive"Hohmann, p. 55 front paws are protected by a thin Stratum corneum|horny layer which becomes pliable when wet.Hohmann, pp. 56–59; MacClintock, p. 15 The five digits of the paws have no webbing between them, which is unusual for a carnivoran .Zeveloff, p. 69 Almost two-thirds of the area responsible for perception|sensory perception in the raccoon's cerebral cortex is specialized for the interpretation of tactile impulses, more than in any other studied animal.Hohmann, p. 56 They are able to identify objects before touching them with vibrissa e located above their sharp, nonretractable claw s.Hohmann, p. 57; Zeveloff, p. 70 The raccoon's paws lack an opposable thumb and thus it does not have the agility of the hands of primate s.MacClintock, p. 15; Zeveloff, p. 70 There is no observed negative effect on tactile perception when a raccoon stands in water below 10& nbsp;°C (50& nbsp;°F) for hours.Hohmann, pp. 60–62

Raccoons are thought to be color blindness|color blind or at least poorly able to distinguish color, though their eyes are well-adapted for sensing green light.Hohmann, p. 63; MacClintock, p. 18; Zeveloff, p. 66 Although their accommodation (eye)|accommodation of 11 dioptre is comparable to that of humans and they see well in twilight because of the tapetum lucidum behind the retina , visual perception is of subordinate importance to raccoons because of their poor long-distance vision.Hohmann, pp. 63–65; MacClintock, pp. 18–21; Zeveloff, pp. 66–67 In addition to being useful for orientation in the dark, their olfaction|sense of smell is important for intraspecific communication. Glandular secretion s (usually from their anal glands ), urine and feces are used for marking.Hohmann, pp. 67–70; MacClintock, p. 17; Zeveloff, pp. 68–69 With their broad hearing (sense)|auditory range , they can perceive tones up to 50–85& nbsp; Hertz|kHz as well as quiet noises like those produced by earthworm s underground.Hohmann, pp. 66, 72; Zeveloff, p. 68

Intelligence

Only a few studies have been undertaken to determine the mental abilities of raccoons, most of them based on the animal's sense of touch. In a study by the ethology|ethologist H. B. Davis in 1908, raccoons were able to open 11 of 13 complex locks in less than 10 tries and had no problems repeating the action when the locks were rearranged or turned upside down. Davis concluded they understood the abstraction|abstract principles of the locking mechanisms and their learning speed was equivalent to that of rhesus macaque s. Studies in 1963, 1973, 1975 and 1992 concentrated on raccoon memory showed they can remember the solutions to tasks for up to three years.Hohmann, pp. 71–72 In a study by B. Pohl in 1992, raccoons were able to instantly differentiate between identical and different symbols three years after the short initial learning phase. Stanislas Dehaene reports in his book The Number Sense raccoons can distinguish boxes containing two or four grapes from those containing three.

Behavior

Social behavior

Studies in the 1990s by the ethologists Stanley D. Gehrt and Ulf Hohmann indicated that raccoons engage in gender-specific social behavior s and are not typically solitary, as was previously thought. (Dissertation at the University of Missouri-Columbia)Hohmann, pp. 133–155 Related females often live in a so-called " fission-fusion society ", that is, they share a common area and occasionally meet at feeding or resting grounds.Bartussek, pp. 10–12; Hohmann, pp. 141–142 Unrelated males often form loose male social groups to maintain their position against foreign males during the mating season —or against other potential invaders.Hohmann, pp. 152–154 Such a group does not usually consist of more than four individuals.Bartussek, p. 12; Hohmann, p. 140 Since some males show aggressive behavior towards unrelated kits, mothers will isolate themselves from other raccoons until their kits are big enough to defend themselves.Hohmann, pp. 124–126, 155 With respect to these three different modes of life prevalent among raccoons, Hohmann called their social structure a "three class society".Hohmann, p. 133 Samuel I. Zeveloff, professor of zoology at Weber State University and author of the book Raccoons: A Natural History , is more cautious in his interpretation and concludes at least the females are solitary most of the time and, according to Erik K. Fritzell's study in North Dakota in 1978, males in areas with low population densities are as well.Zeveloff, pp. 137–139

The shape and size of a raccoon's home range varies depending on age, gender, and habitat, with adults claiming areas more than twice as large as juveniles.MacClintock, p. 61 While the size of home ranges in the inhospitable habitat of North Dakota's prairie s lay between for males and between for females, the average size in a marsh at Lake Erie was .MacClintock, pp. 60–61 Irrespective of whether the home ranges of adjacent groups overlap, they are most likely not actively defended outside the mating season if food supplies are sufficient.Zeveloff, pp. 137–138 Odor marks on prominent spots are assumed to establish home ranges and identify individuals.Zeveloff, pp. 68–69 Urine and feces left at shared latrines may provide additional information about feeding grounds, since raccoons were observed to meet there later for collective eating, sleeping and playing.Hohmann, pp. 142–147

Concerning the general behavior patterns of raccoons, Gehrt points out "typically you'll find 10 to 15 percent that will do the opposite" of what is expected.

Diet

Though usually nocturnal, the raccoon is sometimes active in daylight to take advantage of available food sources.Bartussek, p. 10; Zeveloff, p. 99 Its diet consists of about 40% invertebrate s, 33% plant material and 27% vertebrate s.Hohmann, p. 82 Since its diet consists of such a variety of different foods, Zeveloff argues the raccoon "may well be one of the world's most omnivorous animals".Zeveloff, p. 102 While its diet in spring and early summer consists mostly of insects, worms, and other animals already available early in the year, it prefers fruits and nuts, such as acorn s and walnuts, which emerge in late summer and autumn, and represent a rich calorie source for building up fat needed for winter.Hohmann, pp. 85–88; MacClintock, pp. 44–45 They eat active or large prey, such as bird s and mammal s, only occasionally, since they prefer prey that is easier to catch, specifically fish and amphibian s.Hohmann, p. 83 Bird nests (eggs and after hatchlings) are frequently preyed on, and small birds are often helpless to prevent the attacking raccoon.. Birdfeedersdirect.com. Retrieved on 2011-09-15. When food is plentiful, raccoons can develop strong individual preferences for specific foods.MacClintock, p. 44 In the northern parts of their range, raccoons go into a winter rest , reducing their activity drastically as long as a permanent snow cover makes searching for food impossible.MacClintock, pp. 108–113

Dousing

Raccoons sample food and other objects with their front paws to examine them and to remove unwanted parts. The tactile sensitivity of their paws is increased if this action is performed underwater, since the water softens the horny layer covering the paws.Hohmann, p. 55; Zeveloff, p. 7 However, the behavior observed in captive raccoons in which they carry their food to a watering hole to "wash" or douse it before eating has not been observed in the wild.Lagoni-Hansen, p. 41; MacClintock, pp. 56–57 Natural history|Naturalist Georges-Louis Leclerc, Comte de Buffon (1707–1788) believed that raccoons do not have adequate salivary gland|saliva production to moisten food, necessitating dousing, but this is certainly incorrect.Holmgren, p. 70; Lagoni-Hansen, p. 41; MacClintock, p. 57; Zeveloff, p. 7 Captive raccoons douse their food more frequently when a watering hole with a layout similar to a stream is not farther away than .MacClintock, p. 57 The widely accepted theory is that dousing is a vacuum activity imitating foraging at shores for aquatic foods.Hohmann, pp. 44–45; Lagoni-Hansen, pp. 41–42; MacClintock, p. 57; Zeveloff, p. 7 This is supported by the observation that such foods are doused more frequently. Cleaning dirty food does not seem to be a reason for "washing". Experts have cast doubt on the veracity of observations of wild raccoons dousing food.Holmgren, p. 22 (pro); Lagoni-Hansen, p. 41 (contra); MacClintock, p. 57 (contra)

Reproduction

Raccoons usually mating|mate in a period triggered by increasing daylight between late January and mid-March.Hohmann, p. 150; MacClintock, p. 81; Zeveloff, p. 122 However, there are large regional differences which are not completely explicable by solar conditions. For example, while raccoons in southern states typically mate later than average, the mating season in Manitoba also peaks later than usual in March and extends until June.Zeveloff, p. 122 During the mating season, males roam their home ranges in search of females in an attempt to court them during the three to four day period when conception is possible. These encounters will often occur at central meeting places.Hohmann, pp. 148–150; Lagoni-Hansen, p. 47; MacClintock, pp. 81–82 Sexual intercourse|Copulation , including foreplay, can last over an hour and is repeated over several nights.Hohmann, pp. 150–151 The weaker members of a male social group also are assumed to get the opportunity to mate, since the stronger ones cannot mate with all available females.Hohmann, pp. 153–154 In a study in southern Texas during the mating seasons from 1990 to 1992, about one third of all females mated with more than one male. If a female does not become pregnancy|pregnant or if she loses her kits early, she will sometimes become fertile again 80 to 140& nbsp;days later.Hohmann, p. 125; Lagoni-Hansen, p. 45; Zeveloff, p. 125

After usually 63 to 65& nbsp;days of gestation (although anywhere from 54 to 70& nbsp;days is possible), a litter (animal)|litter of typically two to five young is born.Hohmann, p. 131; Zeveloff, pp. 121, 126 The average litter size varies widely with habitat, ranging from 2.5 in Alabama to 4.8 in North Dakota .Lagoni-Hansen, p. 50; Zeveloff, p. 126 Larger litters are more common in areas with a high mortality rate, due, for example, to hunting or severe winters.Bartussek, p. 32; Zeveloff, p. 126 While male yearlings usually reach their sexual maturity only after the main mating season, female yearlings can compensate for high mortality rates and may be responsible for about 50% of all young born in a year.Hohmann, p. 163; MacClintock, p. 82; Zeveloff, pp. 123–127 Males have no part in raising young.Bartussek, p. 12; Hohmann, p. 111; MacClintock, p. 83 The kits (also called "cubs") are blind and deaf at birth, but their mask is already visible against their light fur.Hohmann, pp. 114, 117; Zeveloff, p. 127 The birth weight of the about -long kits is between .Zeveloff, p. 127 Their ear canals open after around 18 to 23& nbsp;days, a few days before their eyes open for the first time.Hohmann, p. 117 Once the kits weigh about , they begin to explore outside the den, consuming solid food for the first time after six to nine& nbsp;weeks.Hohmann, p. 119; MacClintock, pp. 94–95 After this point, their mother Lactation|suckles them with decreasing frequency; they are usually weaned by 16& nbsp;weeks.Zeveloff, p. 129 In the fall, after their mother has shown them dens and feeding grounds, the juvenile group splits up.Hohmann, pp. 126–127. Zeveloff, p. 130 While many females will stay close to the home range of their mother, males can sometimes move more than away.Hohmann, p. 130; Zeveloff, pp. 132–133 This is considered an instinct ive behavior, preventing inbreeding .Hohmann, p. 128; Zeveloff, p. 133 However, mother and offspring may share a den during the first winter in cold areas.Zeveloff, p. 130

Life expectancy

Captive raccoons have been known to live for more than 20& nbsp;years.Bartussek, p. 6 However, the species' life expectancy in the wild is only 1.8 to 3.1& nbsp;years, depending on the local conditions in terms of traffic volume, hunting, and weather severity.Zeveloff, pp. 118–119 It is not unusual for only half of the young born in one year to survive a full year.Hohmann, p. 163; Zeveloff, p. 119 After this point, the annual mortality rate drops to between 10% and 30%.Hohmann, p. 163 Young raccoons are vulnerable to losing their mother and to starvation, particularly in long and cold winters.MacClintock, p. 73 The most frequent natural cause of death in the North American raccoon population is Canine distemper|distemper , which can reach epidemic proportions and kill most of a local raccoon population. In areas with heavy vehicular traffic and extensive hunting, these factors can account for up to 90% of all deaths of adult raccoons.Hohmann, p. 162

The most important natural predation|predators of the raccoon are bobcat s, coyote s, and great horned owl s, the latter mainly preying on young raccoons. In the Chesapeake Bay , raccoons are the most important mammalian prey for bald eagle s. In their introduced range in the former Soviet Union, their main predators are wolf|wolves , Eurasian lynx|lynx es and eagle owl s. However, predation is not a significant cause of death, especially because larger predators have been Holocene extinction event|exterminated in many areas inhabited by raccoons.Zeveloff, pp. 111–112

Range

Habitat

Although they have thrived in sparsely wooded areas in the last decades, raccoons depend on vertical structures to climb when they feel threatened.Hohmann, pp. 93–94; Zeveloff, p. 93 Therefore, they avoid open terrain and areas with high concentrations of beech|beech trees , as beech bark is too smooth to climb.Hohmann, p. 94 Tree hollow s in old oak s or other trees and rock crevices are preferred by raccoons as sleeping, winter and litter dens. If such dens are unavailable or accessing them is inconvenient, raccoons use burrow s dug by other mammals, dense undergrowth , roadside culverts in urban areas, or tree crotches.Hohmann, pp. 97–101; Zeveloff, pp. 95–96 In a study in the Solling range of hills in Germany, more than 60% of all sleeping places were used only once, but those used at least ten times accounted for about 70% of all uses.Hohmann, p. 98 Since amphibians, crustacean s, and other animals found around the shore of lakes and rivers are an important part of the raccoon's diet, lowland deciduous or Temperate broadleaf and mixed forests|mixed forests abundant with water and marshes sustain the highest population densities.Hohmann, p. 160; Zeveloff, p. 98 While population densities range from 0.5 to 3.2 animals per square kilometre (0.2 – 1.2 animals per square mile) in prairies and do not usually exceed 6 animals per square kilometer (2.3 animals per square mile) in upland hardwood forests, more than 20 raccoons per square kilometer (50 animals per square mile) can live in lowland forests and marshes.Hohmann, p. 160; Zeveloff, p. 97

Distribution in North America

There is evidence that in pre-Columbian times raccoons were numerous only along rivers and in the woodlands of the Southeastern United States .Zeveloff, p. 77 As raccoons were not mentioned in earlier reports of American pioneer|pioneers exploring the central and north-central parts of the United States,Zeveloff, p. 78 their initial spread may have begun a few decades before the 20th century. Since the 1950s, raccoons have expanded their range from Vancouver Island —formerly the northernmost limit of their range—far into the northern portions of the four south-central Canadian provinces.Zeveloff, p. 75 New habitats which have recently been occupied by raccoons (aside from urban areas) include mountain ranges, such as the Western Rocky Mountains , prairies and Salt marsh|coastal marshes .Zeveloff, p. 76 After a population explosion starting in the 1940s, the estimated number of raccoons in North America in the late 1980s was 15 to 20& nbsp;times higher than in the 1930s, when raccoons were comparatively rare.Zeveloff, pp. 75–76 Urbanization , the expansion of agriculture , deliberate introductions, and the extermination of natural predators of the raccoon have probably caused this increase in abundance and distribution.Zeveloff, pp. 76–78

Distribution outside North America

As a result of escapes and deliberate introduced species|introductions in the mid-20th century, the raccoon is now distributed in several European and Asian countries. Sightings have occurred in all the countries bordering Germany, which hosts the largest population outside of North America.Lagoni-Hansen, pp. 89–90 Another stable population exists in northern France, where several pet raccoons were released by members of the United States Air Force|U.S. Air Force near the Laon-Couvron Air Base in 1966.Hohmann, p. 13 About 1,240 animals were released in nine regions of the former Soviet Union between 1936 and 1958 for the purpose of establishing a population to be hunted for their fur. Two of these introductions were successful: one in the south of Belarus between 1954 and 1958, and another in Azerbaijan between 1941 and 1957. With a seasonal harvest of between 1,000 and 1,500& nbsp;animals, in 1974 the estimated size of the population distributed in the Caucasus region was around 20,000& nbsp;animals and the density was four animals per square kilometer (10& nbsp;animals per square mile).Lagoni-Hansen, pp. 90–92

Distribution in Germany

On April 12, 1934, two pairs of pet raccoons were released into the German countryside at the Edersee reservoir in the north of Hesse by forest superintendent Wilhelm Freiherr Sittich von Berlepsch, upon request of their owner, the poultry farmer Rolf Haag.Hohmann, pp. 9–10 He released them two weeks before receiving permission from the Prussia n hunting office to "enrich the fauna ", as Haag's request stated.Hohmann, p. 10 Several prior attempts to introduce raccoons in Germany were not successful.Hohmann, p. 11; Lagoni-Hansen, p. 84 A second population was established in East Germany in 1945 when 25 raccoons escaped from a fur farming|fur farm at Wolfshagen east of Berlin after an air strike. The two populations are parasitologically distinguishable: 70% of the raccoons of the Hessian population are infected with the Nematode|roundworm Baylisascaris procyonis , but none of the Brandenburg ian population has the parasite.Hohmann, p. 182 The estimated number of raccoons was 285 animals in the Hessian region in 1956, over 20,000 animals in the Hessian region in 1970 and between 200,000 and 400,000& nbsp;animals in the whole of Germany in 2008.Hohmann, p. 11

The raccoon was a protected species in Germany, but has been declared a game (food)|game animal in 14 States of Germany|states since 1954.Hohmann, pp. 18, 21 Hunters and environmentalism|environmentalists argue the raccoon spreads uncontrollably, threatens protected bird species and supersedes domestic carnivora ns. This view is opposed by the zoologist Frank-Uwe Michler, who finds no evidence a high population density of raccoons has negative effects on the biodiversity of an area. Hohmann holds extensive hunting cannot be justified by the absence of natural predators, because predation is not a significant cause of death in the North American raccoon population.Hohmann, pp. 14–16

Distribution in the former USSR

Experiments in acclimatising raccoons into the USSR began in 1936, and were repeated a further 25 times until 1962. Overall, 1,222 individuals were released, 64 of which came from zoos and fur farms (38 of them having been imports from western Europe). The remainder originated from a population previously established in Transcaucasia . The range of Soviet raccoons was never single or continuous, as they were often introduced to different locations far from each other. All introductions into the Russian Far East failed ; melanistic raccoons were released on Petrov Island near Vladivostok and some areas of southern Primorsky Krai , but died. In Middle Asia , raccoons were released in Kyrgyzstan 's Jalal-Abad Province , though they were later recorded as "practically absent" there in January 1963. A large and stable raccoon population (yielding 1000–1500 catches a year) was established in Azerbaijan after an introduction to the area in 1937. Raccoons apparently survived an introduction near Terek River|Terek , along the Sulak River into the Dagestan i lowlands. Attempts to settle racoons on the Kuban River 's left tributary and Kabardino-Balkaria were unsuccessful. A successful acclimatization occurred in Belarus , where three introductions (consisting of 52, 37 and 38 individuals in 1954 and 1958) took place. By January 1, 1963, 700 individuals were recorded in the country.

Distribution in Japan

In Japan, up to 1,500& nbsp;raccoons were imported as pets each year after the success of the anime series Rascal the Raccoon (1977). In 2004, the descendants of discarded or escaped animals lived in 42 of 47& nbsp; Prefectures of Japan|prefectures . In Japan, it competes as a wild animal alongside the indigenous raccoon dog (tanuki), a wild canid with a similar lifestyle and visual characteristics.

Urban raccoons

Due to its adaptability, the raccoon has been able to use urban area s as a habitat. The first sightings were recorded in a suburb of Cincinnati in the 1920s. Since the 1950s, raccoons have been present in Philadelphia , Washington, DC , New York City , Chicago , San Francisco , Los Angeles , and Toronto . (Diploma thesis at the University of Halle-Wittenberg) Since the 1960s, Kassel has hosted Europe's first and densest population in a large urban area, with about 50 to 150& nbsp;animals per square kilometre (130–400& nbsp;animals per square mile), a figure comparable to those of urban habitats in North America.Hohmann, p. 108 Home range sizes of urban raccoons are only three to 40 hectares (7.5–100 acres) for females and eight to 80 hectares (20–200 acres) for males. In small towns and suburbs, many raccoons sleep in a nearby forest after foraging in the settlement area.Bartussek, p. 20 Fruit and insects in gardens and leftovers in municipal waste are easily available food sources.Bartussek, p. 21 Furthermore, a large number of additional sleeping areas exist in these areas, such as hollows in old garden trees, cottages, garages, abandoned houses, and attics. The percentage of urban raccoons sleeping in abandoned or occupied houses varies from 15% in Washington, DC (1991) to 43% in Kassel (2003).Bartussek, p. 20; Hohmann, p. 108

Health

Raccoons can carry rabies , a lethal disease caused by the neurotropic virus|neurotropic rabies virus carried in the saliva and transmitted by bites. Its spread began in Florida and Georgia (U.S. state)|Georgia in the 1950s and was facilitated by the introduction of infected individuals to Virginia and North Dakota in the late 1970s.Zeveloff, p. 113 Of the 6,940& nbsp;documented rabies cases reported in the United States in 2006, 2,615 (37.7%) were in raccoons. The United States Department of Agriculture|U.S. Department of Agriculture , as well as local authorities in several U.S. states and Canadian provinces, has developed oral vaccination programs to fight the spread of the disease in endangered populations. Only one human fatality has been reported after transmission of the rabies virus from a raccoon. Among the main symptoms for rabies in raccoons are a generally sickly appearance, impaired mobility, abnormal vocalization, and aggression|aggressiveness . There may be no visible signs at all, however, and most individuals do not show the aggressive behavior seen in infected canids; rabid raccoons will often retire to their dens instead. Organizations like the United States Forest Service|U.S. Forest Service encourage people to stay away from animals with unusual behavior or appearance, and to notify the proper authorities, such as an animal control officer from the local health department . Since healthy animals, especially nursing mothers, will occasionally forage during the day, daylight activity is not a reliable indicator of illness in raccoons.

Unlike rabies and at least a dozen other pathogen s carried by raccoons, Canine distemper|distemper , an epizootic virus, does not affect humans.MacClintock, p. 72; Zeveloff, p. 114 This disease is the most frequent natural cause of death in the North American raccoon population and affects individuals of all age groups. For example, 94 of 145 raccoons died during an outbreak in Clifton, Ohio , in 1968.Zeveloff, p. 112 It may occur along with a following inflammation of the brain ( encephalitis ), causing the animal to display rabies-like symptoms. In Germany, the first eight cases of distemper were reported in 2007.

Some of the most important bacteria l diseases which affect raccoons are leptospirosis , listeriosis , tetanus , and tularemia . Although internal parasitism|parasites weaken their immune system s, well-fed individuals can carry a great many roundworms in their Gastrointestinal tract|digestive tract s without showing symptoms.MacClintock, pp. 73–74; Zeveloff, p. 114 The larvae of the Baylisascaris procyonis roundworm, which can be contained in the feces and seldom causes a severe illness in humans, can be ingested when cleaning raccoon latrines without wearing breathing protection.Hohmann, pp. 169, 182

Raccoons and people

Conflicts

The increasing number of raccoons in urban areas has resulted in diverse reactions in humans, ranging from outrage at their presence to deliberate feeding.Hohmann, pp. 103–106 Some wildlife experts and most public authorities caution against feeding wild animals because they might become increasingly obtrusive and dependent on humans as a food source.Bartussek, p. 34 Other experts challenge such arguments and give advice on feeding raccoons and other wildlife in their books.Holmgren, pp. 117–121 Raccoons without a fear of humans are a concern to those who attribute this trait to rabies, but scientists point out this behavior is much more likely to be a behavioral adjustment to living in habitats with regular contact to humans for many generations.Bartussek, p. 24; Hohmann, p. 182 Serious attacks on humans by groups of non-rabid raccoons are extremely rare and are almost always the result of the raccoon feeling threatened; at least one such attack has been documented. Raccoons usually do not prey on domestic cats and dogs, but individual cases of killings have been reported.

While overturned waste containers and raided fruit trees are just a nuisance to homeowners, it can cost several thousand dollars to repair damage caused by the use of attic space as dens. (Diploma thesis at the University of Halle-Wittenberg) Relocating or killing raccoons without a permit is forbidden in many urban areas on grounds of animal welfare . These methods usually only solve problems with particularly wild or aggressive individuals, since adequate dens are either known to several raccoons or will quickly be rediscovered.Bartussek, p. 32; Hohmann, pp. 142–144, 169 Loud noises, flashing lights and unpleasant odors have proven particularly effective in driving away a mother and her kits before they would normally leave the nesting place (when the kits are about eight weeks old).Bartussek, p. 40 Typically, though, only precautionary measures to restrict access to food waste and denning sites are effective in the long term.Bartussek, pp. 36–40; Hohmann, p. 169

Among all fruits and crops cultivated in agricultural areas, sweet corn in its milk stage is particularly popular among raccoons.Hohmann, pp. 87–88; MacClintock, p 49–50 In a two-year study by Purdue University researchers, published in 2004, raccoons were responsible for 87% of the damage to corn plants. Like other predators, raccoons searching for food can break into poultry houses to feed on chickens, ducks, their eggs, or feed.Hohmann, p. 82; MacClintock, pp. 47–48 Since they may enter tents and try to open locked containers on camping|camping grounds , campers are advised to not keep food or toothpaste inside a tent.

Since raccoons in high mortality areas have a higher rate of reproduction, extensive hunting may not solve problems with raccoon populations. Older males also claim larger home ranges than younger ones, resulting in a lower population density. The costs of large-scale measures to eradicate raccoons from a given area are usually many times higher than the costs of the damage done by the raccoons.

Mythology, arts, and entertainment

In the mythology of the indigenous peoples of the Americas, the raccoon was the subject of folk tale s.Holmgren, pp. 25–46 Stories such as "How raccoons catch so many crayfish " from the Tuscarora (tribe)|Tuscarora centered on its skills at foraging.Holmgren, pp. 41–43 In other tales, the raccoon played the role of the trickster which outsmarts other animals, like coyotes and wolves.Holmgren, pp. 26–29, 38–40 Among others, the Dakota Sioux believed the raccoon had natural spirit power s, since its mask resembled the facial paintings, two-fingered swashes of black and white, used during ritual s to connect to spirit beings.Holmgren, pp. 15–17 The Aztecs linked supernatural abilities especially to females, whose commitment to their young was associated with the role of wise women in the tribal society.Holmgren, pp. 17–18

The raccoon also appears in Native American art across a wide geographic range. Petroglyph s with engraved raccoon tracks were found in Lewis Canyon , Texas; at the Crow Hollow petroglyph site in Grayson County, Kentucky ;Rock Art of Kentucky. Fred E. Coy, Thomas C. Fuller, Larry G. Meadows, James L. Swauger University Press of Kentucky, 2003 p. 60 and Fig. 65APamela LeBlanc . American Statesman. December 13, 2008 and in river drainages near Tularosa , New Mexico and San Francisco , California.Schaafsma, P. Indian Rock Art of the Southwest Albuq., U.NM, 1992 A true-to-detail figurine made of quartz , the ''Ohio Mound Builders' Stone Pipe , was found near the Scioto River . The meaning and significance of the Raccoon Priests Gorget , which features a stylized carving of a raccoon and was found at the Spiro Mounds , Oklahoma, remains unknown.Holmgren, p. 45

In Western culture, several autobiography|autobiographical novel s about living with a raccoon have been written, mostly for Children's literature|children . The best-known is Sterling North 's Rascal (book)|Rascal , which recounts how he raised a kit during World War I . In recent years, anthropomorphism|anthropomorphic raccoons played main roles in the animated television series The Raccoons , the computer-animated film Over the Hedge (film)|Over the Hedge and the video game series Sly Cooper (series)|Sly Cooper .

Hunting and fur trade

The fur of raccoons is used for clothing, especially for Raccoon coat|coats and coonskin cap s. At present, it is the material used for the inaccurately named "sealskin" cap worn by the Royal Fusiliers of Great Britain .A Dictionary of Military Uniform: W.Y.Carman ISBN 0-684-15130-8 Historically, Indigenous people of the Americas|Native American tribes not only used the fur for winter clothing, but also used the tails for ornament.Holmgren, p. 18 Since the late 18th century, various types of scent hound s which are able to tree animals (" coonhound s") have been bred in the United States. In the 19th century, when coonskins occasionally even served as means of payment, several thousand raccoons were killed each year in the United States.Holmgren, p. 74; Zeveloff, p. 160 This number rose quickly when automobile coats became popular after the turn of the 20th century. In the 1920s, wearing a raccoon coat was regarded as status symbol among college student s.Holmgren, p. 77 Attempts to breed raccoons in fur farms in the 1920s and 1930s in North America and Europe turned out not to be profitable, and farming was abandoned after prices for long-haired pelts dropped in the 1940s.Zeveloff, p. 161 Although raccoons had become rare in the 1930s, at least 388,000 were killed during the hunting season of 1934/35.Holmgren, p. 77; Zeveloff, pp. 75, 160, 173

After persistent population increases began in the 1940s, the seasonal hunt reached about one million animals in 1946/47 and two million in 1962/63.Zeveloff, pp. 75, 160 The 1948 senatorial campaign of Estes Kefauver , who wore such a cap for promotional purposes,Fontenay, Charles L. Estes Kefauver: A Biography. TN, 1980. rev by Salvatore LaGumina, Annals of the American Academy of Political and Social Science, 462 (1982), p.180 and the broadcast of three television episodes about the Frontier#American frontier|frontiersman Davy Crockett and the film Davy Crockett, King of the Wild Frontier in 1954 and 1955 led to a high demand for coonskin cap s in the United States (though the caps supplied to the fad were typically made of faux fur with a raccoon tail attached). Ironically, it is unlikely either Crockett or the actor who played him, Fess Parker , actually wore a cap made from raccoon fur.Zeveloff, p. 170 The seasonal hunt reached an all-time high with 5.2& nbsp;million animals in 1976/77 and ranged between 3.2 and 4.7& nbsp;million for most of the 1980s. In 1982, the average pelt price was $20.The Red Panda, Olingos, Coatis, Raccoons, and Their Relatives: Status Survey and Conservation Action Plan for Procyonids and Ailurids By A. R. Glatston, IUCN/SSC Mustelid, Viverrid & Procyonid Specialist Group Edition: illustrated Published by IUCN, 1994, p. 9 ISBN 2-8317-0046-9, ISBN 978-2-8317-0046-5 In the first half of the 1990s, the seasonal hunt dropped to 0.9 to 1.9& nbsp;million due to decreasing pelt prices.Zeveloff, pp. 160–161 As of 1987, the raccoon was identified as the most important wild furbearer in North America in terms of revenue.The Red Panda, Olingos, Coatis, Raccoons, and Their Relatives: Status Survey and Conservation Action Plan for Procyonids and Ailurids By A. R. Glatston, IUCN/SSC Mustelid, Viverrid & Procyonid Specialist Group Published by IUCN, 1994, p. 9

In many parts of the United States, raccoon hunting is still done at night with dogs, usually breeds of coonhounds. The dogs tracking (dog)|track the raccoon until it seeks refuge, usually in a tree, where it is either harvested or left for future hunts. Hunters can tell the progress of tracking by the type of bark emitted by the dogs; a unique bark indicates the raccoon has been " treeing|treed ".

As food

While primarily hunted for their fur, raccoons were also a source of food for Native Americans and AmericansHolmgren, pp. 18–19, Zeveloff, p. 165 and barbecued raccoon was a traditional food on American farms.Farm: A Year in the Life of an American Farmer. Richard Rhodes, reprint, U of Nebraska Press, 1997, p.270. It was often a festive meal. Raccoon was eaten by Slavery in the United States|American slaves at Christmas ,Harriet Jacobs, Incidents in the Life of a Slave Girl, Digireads.com Publishing, 2005, p.72. but it was not necessarily a dish of the poor or rural; in San Francisco's The Golden Era of December 21, 1856, raccoon is among the specialties advertised for the holiday, and US President Calvin Coolidge 's pet raccoon Rebecca was originally sent to be served at the White House Thanksgiving dinner|Thanksgiving Dinner .San Diego's Hilarious History By Herbert Lockwood, William Carroll Published by Coda Publications, 2004, p. 46.Jen O'Neill. . findingdulcinea.com. November 12, 2008. . The first edition of The Joy of Cooking , released in 1931, contained a recipe for preparing raccoon.

Because raccoons are generally thought of as endearing, cute, and/or varmints , the idea of eating them is repulsive to mainstream consumers. However, many thousands of raccoons are still eaten each year in the United States. Although the Delafield, Wisconsin|Delafield (Wisconsin) Coon Feed has been an annual event since 1928, its culinary use is mainly identified with certain regions of the Southern United States|American South like Arkansas where the Gillett, Arkansas|Gillett Coon Supper is an important political event.

As pets

As with most exotic pets, owning a raccoon often takes a significant amount of time and patience. Raccoons may act unpredictably and aggressively and it can be quite difficult to teach them to obey and understand commands.Bartussek, p. 44; Hohmann, pp. 173–174 In places where keeping raccoons as pets is not forbidden, such as in Wisconsin and other U.S. states, an exotic pet permit may be required.MacClintock, p. 129

Sexually mature raccoons often show aggressive natural behaviors such as biting during the mating season.Bartussek, p. 44; Hohmann, pp. 185–186 Neutering them at around five or six months of age decreases the chances of aggressive behavior developing.Hohmann, p. 186 Raccoons can become obesity|obese and suffer from other disorders due to poor diet and lack of exercise.Hohmann, p. 185 When fed with cat food over a long time period, raccoons can develop gout .Hohmann, p. 180 With respect to the research results regarding their social behavior, it is now required by law in Austria and Germany to keep at least two individuals to prevent loneliness. Raccoons are usually kept in a pen (enclosure)|pen (indoor or outdoor), also a legal requirement in Austria and Germany, rather than in the apartment where their natural curiosity may result in damage to property.Bartussek, p. 44; Hohmann, pp. 184, 187; MacClintock, p. 130–131

When orphaned, it is possible for kits to be wildlife rehabilitation|rehabilitated and Reintroduction|reintroduced to the wild. However, it is uncertain whether they readapt well to life in the wild.MacClintock, p. 130 Feeding unweaned kits with cow's milk rather than a kitten replacement milk or a similar product can be dangerous to their health.Bartussek, p. 44; Hohmann, pp. 175–176

Notes

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